Winner – 15 to 17 years old, and Young Grand Title winner: The fox that got the goose by Liina Heikkinen, Finland
It was on a summer holiday in Helsinki that Liina, then aged 13, heard about a large fox family living in the city suburbs on the island of Lehtisaari. The island has both wooded areas and fox-friendly citizens, and the foxes are relatively unafraid of humans. So Liina spent one long July day, without a hide, watching the two adults and their six large cubs.
Winner – Behaviour, Invertebrates: A tale of two wasps by Frank Deschandol, France
This remarkable simultaneous framing of a red-banded sand wasp (left) and a cuckoo wasp, about to enter next-door nest holes in Normandy, France. Though these two species don’t regularly interact, Deschandol was gifted a perfectly balanced composition by the insects’ fortuitous flight paths to their nest holes.
Winner – Animal Portraits: The pose by Mogens Trolle, Denmark
A young male proboscis monkey cocks his head slightly and closes his eyes. Unexpected pale blue eyelids now complement his immaculately groomed auburn hair. He poses for a few seconds as if in meditation. He is a wild visitor to the feeding station at Labuk Bay Proboscis Monkey Sanctuary in Sabah, Borneo – ‘the most laid-back character,’ says Trolle, ‘quite unlike anything I’ve ever seen on another monkey’ – connects us, he hopes, with a fellow primate.